Pierre Sage endured a frustrating return to Lyon in the Europa League as his Crystal Palace side lost 3-2 to his former employers at the Groupama Stadium.

Corentin Tolisso opened the scoring for the hosts and Ernest Nuamah doubled Lyon’s advantage late in a first half without a shot on target for the visitors. Yéremy Pino halved the deficit five minutes after the break before Pavel Sulc restored the Ligue 1 side’s two-goal advantage. The Palace substitute Daichi Kamada pulled one back with nine minutes remaining in a better second half for Palace, who could not find an equaliser.

Days after beating his predecessor, the Nottingham Forest manager, Oliver Glasner, at Selhurst Park in the Premier League on Sunday, Sage found himself returning to the home of the club he managed for 14 months.

Ismaïla Sarr, who claimed the Conference League golden boot last season, was in the Crystal Palace starting lineup for the first time this season.

Celtic found clinical Benfica just too much to handle during a 4-1 defeat in Lisbon.

Colombia striker Jhon Duran scored for the hosts with a penalty after just five minutes after being fouled by Dane Murray and then helped himself to a second in the 35th minute, after defender Alexander Bah had added a second midway through the first half.

Martin O’Neill’s side were cut apart at times but could have scored themselves on a couple of occasions before the break.

The visitors eventually reduced the deficit in the 63rd minute through a brave header from Camilo Durán – who had joined up with the squad late after remaining in Glasgow for the birth of his second child – but they could not close the gap any further before substitute Vangelis Pavlidis fired in a fourth for the home side with two minutes remaining.

Randal Kolo Muani scored a stoppage-time header to give Juventus a 2-1 win at Celta Vigo which leaves them top of the standings.

Hugo Álvarez had put the home side in front early on before Nick Woltemade equalised on the hour and Kolo Muani then nodded in from close range during the second minute of added time.

Milan cruised to a 4-0 victory at RB Salzburg, courtesy of a hat-trick by the Italian teenager Alphadjo Cissè.

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Milan’s record signing Gonçalo Ramos made it four on 72 minutes before a red card for Filippo Terracciano took some of the shine off the win for Ruben Amorim’s side.

Bayer Leverkusen put themselves top of the standings after the early kick-offs as they followed up their opening 2-0 win with a 4-0 success at Polish side Lech Poznan. The Czech striker Patrik Schick was on target twice from the penalty spot in the first half, either side of a Malik Tillman goal, to ensure the points were wrapped up for Carles Martínez’s side before the interval. Ibrahim Maza added a late fourth.

Real Sociedad followed up their opening defeat at home to Bournemouth with a 1-1 draw at Union Saint-Gilloise.